I tried loading your save in the latest experimental (2.0.19) and got exactly the same crash.
I was expecting it to be something related to migrating circuit signals with quality, but the line that crashed is C:\Users\build\AppData\Local\Temp\factorio-build-qKZ8vt\src\Entity\Beacon.cpp(161) So ...
I can open the constant combinator, and add a new signal but when i delete the old one Factorio crash.
When i build a new constant combinator, Factorio crash when i wire it with the old circuit network
When i turn on/off or deconstruct a old combinator(no matter which one), the Factorio Game crash.
Building a new red wire to the old logic Factorio also crash.